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STREME - Sensitive, Thorough, Rapid, Enriched Motif Elicitation
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MEME version 5.4.1 (Release date: Sat Aug 21 19:23:23 2021 -0700)

For further information on how to interpret these results please access https://meme-suite.org/meme.
To get a copy of the MEME Suite software please access https://meme-suite.org.

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REFERENCE
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If you use this program in your research, please cite:

Timothy L. Bailey,
"STREME: accurate and versatile sequence motif discovery",
Bioinformatics, Mar. 24, 2021.
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ALPHABET= ACGT

strands: + -

Background letter frequencies
A 0.255 C 0.245 G 0.245 T 0.255 

MOTIF 1-VTGASTCAB STREME-1
letter-probability matrix: alength= 4 w= 9 nsites= 3309 E= 3.0e-045
 0.403563 0.259568 0.293430 0.043439
 0.000026 0.007266 0.000720 0.991989
 0.000756 0.000025 0.865417 0.133802
 0.977024 0.014961 0.000025 0.007990
 0.097722 0.402278 0.402278 0.097722
 0.007990 0.000025 0.014961 0.977024
 0.133802 0.865417 0.000025 0.000756
 0.991989 0.000720 0.007266 0.000026
 0.043439 0.293430 0.259568 0.403563

MOTIF 2-TTGMKCAA STREME-2
letter-probability matrix: alength= 4 w= 8 nsites= 484 E= 4.2e-006
 0.051593 0.000379 0.000379 0.947649
 0.000395 0.000379 0.000379 0.998847
 0.198247 0.000379 0.800979 0.000395
 0.294736 0.505873 0.078100 0.121291
 0.121291 0.078100 0.505873 0.294736
 0.000395 0.800979 0.000379 0.198247
 0.998847 0.000379 0.000379 0.000395
 0.947649 0.000379 0.000379 0.051593

MOTIF 3-ACATTCCW STREME-3
letter-probability matrix: alength= 4 w= 8 nsites= 1225 E= 9.6e-006
 0.999170 0.000273 0.000273 0.000284
 0.000284 0.797746 0.201686 0.000284
 0.980466 0.000273 0.018977 0.000284
 0.000284 0.000273 0.000273 0.999170
 0.287346 0.000273 0.000273 0.712108
 0.000284 0.999159 0.000273 0.000284
 0.000284 0.804337 0.000273 0.195106
 0.452234 0.000273 0.011177 0.536317

MOTIF 4-CCCCRCCCC STREME-4
letter-probability matrix: alength= 4 w= 9 nsites= 1811 E= 7.3e-005
 0.007419 0.969330 0.000084 0.023167
 0.006995 0.867723 0.000483 0.124799
 0.153418 0.804062 0.010928 0.031592
 0.048201 0.908512 0.038963 0.004325
 0.543121 0.000084 0.424260 0.032535
 0.012461 0.964028 0.000084 0.023427
 0.004629 0.983762 0.011522 0.000087
 0.007355 0.931714 0.011763 0.049168
 0.177654 0.599402 0.007431 0.215513

MOTIF 5-GATGACDHA STREME-5
letter-probability matrix: alength= 4 w= 9 nsites= 2492 E= 9.8e-005
 0.041792 0.048911 0.831436 0.077861
 0.818009 0.000056 0.166524 0.015411
 0.000058 0.062264 0.000056 0.937623
 0.020479 0.000056 0.833630 0.145836
 0.913191 0.000056 0.009951 0.076802
 0.059732 0.790867 0.047421 0.101980
 0.444148 0.010289 0.319911 0.225652
 0.328832 0.404291 0.038454 0.228424
 0.654753 0.164679 0.114193 0.066375

MOTIF 6-AGAAACA STREME-6
letter-probability matrix: alength= 4 w= 7 nsites= 2492 E= 8.4e-004
 0.998614 0.000456 0.000456 0.000475
 0.000475 0.219730 0.585024 0.194771
 0.998614 0.000456 0.000456 0.000475
 0.998614 0.000456 0.000456 0.000475
 0.959000 0.040070 0.000456 0.000475
 0.000475 0.729478 0.000456 0.269591
 0.684835 0.314235 0.000456 0.000475

MOTIF 7-TGASTCA STREME-7
letter-probability matrix: alength= 4 w= 7 nsites= 767 E= 6.8e-003
 0.000658 0.139348 0.000632 0.859363
 0.000658 0.000632 0.852019 0.146691
 0.998078 0.000632 0.000632 0.000658
 0.000658 0.499342 0.499342 0.000658
 0.000658 0.000632 0.000632 0.998078
 0.146691 0.852019 0.000632 0.000658
 0.859363 0.000632 0.139348 0.000658

MOTIF 8-AGGAAR STREME-8
letter-probability matrix: alength= 4 w= 6 nsites= 9272 E= 2.9e-002
 0.738409 0.000189 0.261205 0.000197
 0.000197 0.000189 0.999417 0.000197
 0.000197 0.111111 0.888495 0.000197
 0.999425 0.000189 0.000189 0.000197
 0.999425 0.000189 0.000189 0.000197
 0.423886 0.000189 0.575728 0.000197

MOTIF 9-CATTCC STREME-9
letter-probability matrix: alength= 4 w= 6 nsites= 1238 E= 7.1e-002
 0.023182 0.931364 0.022273 0.023182
 0.932273 0.022273 0.022273 0.023182
 0.023182 0.022273 0.022273 0.932273
 0.023182 0.022273 0.022273 0.932273
 0.023182 0.931364 0.022273 0.023182
 0.023182 0.931364 0.022273 0.023182

MOTIF 10-CTTGGCAA STREME-10
letter-probability matrix: alength= 4 w= 8 nsites= 849 E= 1.2e-001
 0.009416 0.851688 0.060898 0.077998
 0.012225 0.000600 0.013601 0.973574
 0.010063 0.013601 0.000600 0.975736
 0.062262 0.029798 0.862890 0.045050
 0.001468 0.000600 0.988586 0.009346
 0.077203 0.862457 0.025176 0.035165
 0.797165 0.161260 0.005714 0.035861
 0.800078 0.120529 0.000600 0.078793

MOTIF 11-AAATRC STREME-11
letter-probability matrix: alength= 4 w= 6 nsites= 1600 E= 2.9e-001
 0.995261 0.001559 0.001559 0.001622
 0.995261 0.001559 0.001559 0.001622
 0.995261 0.001559 0.001559 0.001622
 0.001622 0.079027 0.001559 0.917793
 0.376968 0.001559 0.619851 0.001622
 0.001622 0.995197 0.001559 0.001622

MOTIF 12-AGTCACCT STREME-12
letter-probability matrix: alength= 4 w= 8 nsites= 428 E= 3.9e-001
 0.911791 0.070412 0.017118 0.000680
 0.015069 0.000653 0.983598 0.000680
 0.000680 0.000653 0.000653 0.998015
 0.032648 0.879531 0.038470 0.049351
 0.932084 0.000653 0.066583 0.000680
 0.002912 0.842641 0.000653 0.153795
 0.040724 0.844407 0.084186 0.030683
 0.092122 0.098980 0.078526 0.730373

MOTIF 13-AAATATTTRTTG STREME-13
letter-probability matrix: alength= 4 w= 12 nsites= 132 E= 4.8e-001
 0.960893 0.028576 0.000250 0.010281
 0.872976 0.035658 0.066922 0.024444
 0.868834 0.021494 0.045679 0.063993
 0.007342 0.049821 0.014413 0.928425
 0.878854 0.014413 0.078147 0.028586
 0.056911 0.127716 0.007332 0.808041
 0.007342 0.059840 0.028576 0.904242
 0.034463 0.007332 0.000250 0.957955
 0.618018 0.038597 0.343125 0.000261
 0.007027 0.042739 0.014413 0.935821
 0.046289 0.000250 0.021494 0.931966
 0.124425 0.021494 0.787149 0.066932

MOTIF 14-CCGCCTCCTC STREME-14
letter-probability matrix: alength= 4 w= 10 nsites= 234 E= 5.0e-001
 0.009099 0.838290 0.094699 0.057912
 0.000670 0.969953 0.028707 0.000670
 0.000670 0.000644 0.998016 0.000670
 0.049453 0.914396 0.031972 0.004179
 0.000670 0.994507 0.004153 0.000670
 0.044663 0.018859 0.078407 0.858070
 0.018885 0.953479 0.005599 0.022037
 0.073536 0.887885 0.019693 0.018885
 0.074960 0.033130 0.084389 0.807520
 0.033665 0.898914 0.045384 0.022037

MOTIF 15-AAACCWGT STREME-15
letter-probability matrix: alength= 4 w= 8 nsites= 396 E= 6.0e-001
 0.996938 0.001007 0.001007 0.001048
 0.996938 0.001007 0.001007 0.001048
 0.996938 0.001007 0.001007 0.001048
 0.001048 0.996897 0.001007 0.001048
 0.001048 0.656136 0.182153 0.160664
 0.344046 0.001007 0.048546 0.606401
 0.001048 0.001007 0.996897 0.001048
 0.001048 0.001007 0.001007 0.996938

MOTIF 16-ATGATGCAATC STREME-16
letter-probability matrix: alength= 4 w= 11 nsites= 202 E= 6.1e-001
 0.745133 0.008060 0.237236 0.009571
 0.000286 0.008060 0.042431 0.949223
 0.151085 0.023285 0.638670 0.186960
 0.704555 0.000275 0.034646 0.260524
 0.015856 0.000275 0.031416 0.952452
 0.117237 0.000275 0.865132 0.017356
 0.000286 0.894718 0.000275 0.104721
 0.916176 0.003506 0.000275 0.080043
 0.999163 0.000275 0.000275 0.000286
 0.047720 0.081358 0.030092 0.840829
 0.270261 0.576504 0.070339 0.082897

MOTIF 17-ACACCCACAT STREME-17
letter-probability matrix: alength= 4 w= 10 nsites= 229 E= 7.3e-001
 0.750556 0.181535 0.036540 0.031369
 0.002014 0.915036 0.009073 0.073877
 0.820634 0.022447 0.133851 0.023068
 0.091537 0.868855 0.008202 0.031406
 0.000164 0.894862 0.063327 0.041647
 0.102695 0.688073 0.061809 0.147422
 0.855422 0.000158 0.133457 0.010963
 0.050361 0.905113 0.014390 0.030136
 0.686570 0.044574 0.208993 0.059863
 0.149407 0.145478 0.012933 0.692182

MOTIF 18-CTGGGA STREME-18
letter-probability matrix: alength= 4 w= 6 nsites= 4743 E= 7.5e-001
 0.008124 0.962657 0.016566 0.012653
 0.000509 0.120529 0.105586 0.773376
 0.003775 0.001321 0.981842 0.013063
 0.000509 0.019629 0.979353 0.000509
 0.000509 0.000489 0.989764 0.009237
 0.651334 0.148942 0.073999 0.125725

MOTIF 19-TGCGCAGGCGCAV STREME-19
letter-probability matrix: alength= 4 w= 13 nsites= 78 E= 8.1e-001
 0.000990 0.216327 0.027873 0.754810
 0.027912 0.000952 0.970146 0.000990
 0.000990 0.970146 0.000952 0.027912
 0.000990 0.000952 0.943224 0.054834
 0.000990 0.997068 0.000952 0.000990
 0.943263 0.027873 0.027873 0.000990
 0.135598 0.108639 0.727851 0.027912
 0.108677 0.027873 0.862460 0.000990
 0.000990 0.997068 0.000952 0.000990
 0.000990 0.000952 0.997068 0.000990
 0.000990 0.943224 0.054795 0.000990
 0.727883 0.027873 0.243254 0.000990
 0.458659 0.243254 0.243254 0.054834

MOTIF 20-CRGCCAATC STREME-20
letter-probability matrix: alength= 4 w= 9 nsites= 73 E= 8.8e-001
 0.001157 0.996576 0.001111 0.001157
 0.520480 0.001111 0.477252 0.001157
 0.132985 0.001111 0.864747 0.001157
 0.001157 0.996576 0.001111 0.001157
 0.001157 0.927644 0.070043 0.001157
 0.926872 0.001111 0.070860 0.001157
 0.996621 0.001111 0.001111 0.001157
 0.001157 0.001111 0.001111 0.996621
 0.001157 0.996576 0.001111 0.001157

MOTIF 21-CACACACACACACA STREME-21
letter-probability matrix: alength= 4 w= 14 nsites= 168 E= 1.1e+000
 0.038293 0.900337 0.000269 0.061101
 0.991580 0.000269 0.007871 0.000280
 0.015485 0.839514 0.000269 0.144732
 0.793909 0.030679 0.137118 0.038293
 0.000280 0.881976 0.071848 0.045896
 0.900348 0.000269 0.023077 0.076307
 0.000280 0.772381 0.082607 0.144733
 0.953567 0.000269 0.000269 0.045896
 0.000280 0.745127 0.117465 0.137129
 0.775548 0.053488 0.071849 0.099115
 0.000280 0.881976 0.018630 0.099115
 0.748293 0.030680 0.091501 0.129527
 0.083910 0.725474 0.000269 0.190347
 0.748293 0.076296 0.099104 0.076307

MOTIF 22-RGGYAGRG STREME-22
letter-probability matrix: alength= 4 w= 8 nsites= 1269 E= 1.2e+000
 0.588086 0.012672 0.398902 0.000339
 0.000339 0.000326 0.998996 0.000339
 0.000339 0.000326 0.998996 0.000339
 0.000339 0.661630 0.000326 0.337705
 0.959483 0.039852 0.000326 0.000339
 0.000339 0.000326 0.998996 0.000339
 0.459338 0.191156 0.349167 0.000339
 0.000339 0.000326 0.998996 0.000339

MOTIF 23-GTGCCTGGCACABAG STREME-23
letter-probability matrix: alength= 4 w= 15 nsites= 77 E= 3.1e+000
 0.221016 0.033770 0.744707 0.000507
 0.000507 0.069396 0.014269 0.915828
 0.096980 0.014269 0.888244 0.000507
 0.028071 0.750427 0.000487 0.221016
 0.000507 0.998499 0.000487 0.000507
 0.014289 0.000487 0.014269 0.970955
 0.110761 0.014269 0.860681 0.014289
 0.069417 0.014269 0.833117 0.083198
 0.138325 0.805553 0.000487 0.055634
 0.929610 0.000487 0.000487 0.069416
 0.000507 0.846899 0.083178 0.069417
 0.902046 0.014269 0.041832 0.041852
 0.124543 0.226716 0.303689 0.345052
 0.874483 0.055614 0.000487 0.069416
 0.110761 0.000487 0.860681 0.028071

MOTIF 24-AATGAATGAATR STREME-24
letter-probability matrix: alength= 4 w= 12 nsites= 123 E= 4.7e+000
 0.961467 0.000332 0.028475 0.009726
 0.784812 0.009713 0.186368 0.019108
 0.028489 0.009713 0.000332 0.961466
 0.051145 0.131671 0.816839 0.000345
 0.998992 0.000332 0.000332 0.000345
 0.816850 0.075383 0.037858 0.069909
 0.047253 0.000332 0.009713 0.942703
 0.094161 0.037856 0.867638 0.000345
 0.896213 0.019095 0.009713 0.074979
 0.914558 0.028476 0.047240 0.009726
 0.143367 0.060513 0.037857 0.758264
 0.565527 0.060513 0.373614 0.000345

MOTIF 25-CCWCCYACK STREME-25
letter-probability matrix: alength= 4 w= 9 nsites= 172 E= 4.8e+000
 0.000688 0.997964 0.000661 0.000688
 0.000688 0.960571 0.038053 0.000688
 0.440498 0.000661 0.000661 0.558180
 0.000688 0.997964 0.000661 0.000688
 0.000688 0.997964 0.000661 0.000688
 0.000688 0.637479 0.029462 0.332371
 0.997991 0.000661 0.000661 0.000688
 0.000688 0.997964 0.000661 0.000688
 0.000688 0.000661 0.348570 0.650081

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Stopped because 3 consecutive motifs exceeded the p-value threshold (0.05).
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COMMAND:	streme --verbosity 1 --oc streme_out -dna --p ./seqs-centered --minw 6 --maxw 15 --time 5667 --order 2 --thresh 0.05 --totallength 4000000
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CPU:		noble-meme.grid.gs.washington.edu
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FINALTIME:	1648.15 seconds
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